Marine Habitats Classification |
 Littoral rock (and other hard substrata) |
  High energy littoral rock |
   Mussel and/or barnacle communities |
    Mytilus edulis and barnacles on very exposed eulittoral rock |
    Chthamalus spp. on exposed eulittoral rock |
     Chthamalus spp. on exposed upper eulittoral rock |
     Chthamalus spp. and Lichina pygmaea on steep exposed upper eulittoral rock |
    Semibalanus balanoides on exposed to moderately exposed or vertical sheltered eulittoral rock |
     Semibalanus balanoides, Patella vulgata and Littorina spp. on exposed to moderately exposed or vertical sheltered eulittoral rock |
     Semibalanus balanoides, Fucus vesiculosus and red seaweeds on exposed to moderately exposed eulittoral rock |
     Semibalanus balanoides and Littorina spp. on exposed to moderately exposed eulittoral boulders and cobbles |
   Robust fucoid and/or red seaweed communities |
    Fucus distichus and Fucus spiralis f. nana on extremely exposed upper shore rock |
    Corallina officinalis on exposed to moderately exposed lower eulittoral rock |
     Corallina officinalis and Mastocarpus stellatus on exposed to moderately exposed lower eulittoral rock |
     Corallina officinalis, Himanthalia elongata and Patella ulyssiponensis on very exposed lower eulittoral rock |
    Himanthalia elongata and red seaweeds on exposed to moderately exposed lower eulittoral rock |
    Palmaria palmata on very exposed to moderately exposed lower eulittoral rock |
    Mastocarpus stellatus and Chondrus crispus on very exposed to moderately exposed lower eulittoral rock |
    Osmundea pinnatifida on moderately exposed mid eulittoral rock |
    Ceramium sp. and piddocks on eulittoral fossilised peat |
   Fucoids in tide-swept conditions |
    Ascophyllum nodosum, sponges and ascidians on tide-swept mid eulittoral rock |
    Fucus serratus, sponges and ascidians on tide-swept lower eulittoral rock |
    Fucus serratus with sponges, ascidians and red seaweeds on tide-swept lower eulittoral mixed substrata |
  Moderate energy littoral rock |
   Mussels and fucoids on moderately exposed shores |
    Mytilus edulis and Fucus vesiculosus on moderately exposed mid eulittoral rock |
    Mytilus edulis, Fucus serratus and red seaweeds on moderately exposed lower eulittoral rock |
    Mytilus edulis and piddocks on eulittoral firm clay |
   Barnacles and fucoids on moderately exposed shores |
    Pelvetia canaliculata and barnacles on moderately exposed littoral fringe rock |
    Fucus spiralis on exposed to moderately exposed upper eulittoral rock |
    Fucus vesiculosus and barnacle mosaics on moderately exposed mid eulittoral rock |
    Fucus serratus on moderately exposed lower eulittoral rock |
     Fucus serratus and red seaweeds on moderately exposed lower eulittoral rock |
     Fucus serratus and under-boulder fauna on exposed to moderately exposed lower eulittoral boulders |
     Fucus serratus and piddocks on lower eulittoral soft rock |
    Rhodothamniella floridula on sand-scoured lower eulittoral rock |
  Low energy littoral rock |
   Fucoids on sheltered marine shores |
    Pelvetia canaliculata on sheltered littoral fringe rock |
    Fucus spiralis on sheltered upper eulittoral rock |
     Fucus spiralis on full salinity sheltered upper eulittoral rock |
     Fucus spiralis on full salinity upper eulittoral mixed substrata |
    Fucus vesiculosus on moderately exposed to sheltered mid eulittoral rock |
     Fucus vesiculosus on full salinity moderately exposed to sheltered mid eulittoral rock |
     Fucus vesiculosus on mid eulittoral mixed substrata |
    Ascophyllum nodosum on very sheltered mid eulittoral rock |
     Ascophyllum nodosum on full salinity mid eulittoral rock |
     Ascophyllum nodosum on full salinity mid eulittoral mixed substrata |
    Fucus serratus on sheltered lower eulittoral rock |
     Fucus serratus on full salinity sheltered lower eulittoral rock |
     Fucus serratus on full salinity lower eulittoral mixed substrata |
   Fucoids in variable salinity |
    Pelvetia canaliculata on sheltered variable salinity littoral fringe rock |
    Fucus spiralis on sheltered variable salinity upper eulittoral rock |
    Fucus vesiculosus on variable salinity mid eulittoral boulders and stable mixed substrata |
    Ascophyllum nodosum and Fucus vesiculosus on variable salinity mid eulittoral rock |
    Ascophyllum nodosum ecad mackaii beds on extremely sheltered mid eulittoral mixed substrata |
    Fucus serratus and large Mytilus edulis on variable salinity lower eulittoral rock |
    Fucus ceranoides on reduced salinity eulittoral rock |
  Features of littoral rock |
   Lichens or small green algae on supralittoral and littoral fringe rock |
    Yellow and grey lichens on supralittoral rock |
    Prasiola stipitata on nitrate-enriched supralittoral or littoral fringe rock |
    Verrucaria maura on littoral fringe rock |
     Verrucaria maura and sparse barnacles on exposed littoral fringe rock |
     Verrucaria maura on very exposed to very sheltered upper littoral fringe rock |
    Blidingia spp. on vertical littoral fringe soft rock |
    Ulothrix flacca and Urospora spp. on freshwater-influenced vertical littoral fringe soft rock |
   Rockpools |
    Green seaweeds (Enteromorpha spp. and Cladophora spp.) in shallow upper shore rockpools |
    Coralline crust-dominated shallow eulittoral rockpools |
     Coralline crusts and Corallina officinalis in shallow eulittoral rockpools |
     Coralline crusts and Paracentrotus lividus in shallow eulittoral rockpools |
     Bifurcaria bifurcata in shallow eulittoral rockpools |
     Cystoseira spp. in eulittoral rockpools |
    Fucoids and kelp in deep eulittoral rockpools |
     Sargassum muticum in eulittoral rockpools |
    Seaweeds in sediment-floored eulittoral rockpools |
    Hydroids, ephemeral seaweeds and Littorina littorea in shallow eulittoral mixed substrata pools |
   Littoral caves and overhangs |
    Chrysophyceae and Haptophyceae on vertical upper littoral fringe soft rock |
    Green algal films on upper and mid-shore cave walls and ceilings |
    Audouinella purpurea and Pilinia maritima crusts on upper and mid-shore cave walls and ceilings |
    Audouinella purpurea and Cladophora rupestris on upper to mid-shore cave walls |
    Verrucaria mucosa and/or Hildenbrandia rubra on upper to mid shore cave walls |
    Sponges and shade-tolerant red seaweeds on overhanging lower eulittoral bedrock and in cave entrances |
     Sponges, shade-tolerant red seaweeds and Dendrodoa grossularia on wave-surged overhanging lower eulittoral bedrock and caves |
    Sponges, bryozoans and ascidians on deeply overhanging lower shore bedrock or caves |
    Faunal crusts on wave-surged littoral cave walls |
    Sparse fauna (barnacles and spirorbids) on sand/pebble-scoured rock in littoral caves |
    Barren and/or boulder-scoured littoral cave walls and floors |
   Ephemeral green or red seaweed communities (freshwater or sand-influenced) |
   Enteromorpha spp. on freshwater-influenced and/or unstable upper eulittoral rock |
   Porphyra purpurea and Enteromorpha spp. on sand-scoured mid or lower eulittoral rock |
   Ephemeral green and red seaweeds on variable salinity and/or disturbed eulittoral mixed substrata |
   Barnacles and Littorina spp. on unstable eulittoral mixed substrata |
 Littoral sediment |
  Littoral coarse sediment |
   Shingle (pebble) and gravel shores |
    Barren littoral shingle |
    Pectenogammarus planicrurus in mid shore well-sorted gravel or coarse sand |
  Littoral sand |
   Strandline |
    Talitrids on the upper shore and strand-line |
    Mytilus edulis and Fabricia sabella in littoral mixed sediment |
   Barren or amphipod-dominated mobile sand shores |
    Barren littoral coarse sand |
    Oligochaetes in littoral mobile sand |
     Oligochaetes in full salinity littoral mobile sand |
     Oligochaetes in variable salinity littoral mobile sand |
    Amphipods and Scolelepis spp. in littoral medium-fine sand |
     Scolelepis spp. in littoral mobile sand |
     Eurydice pulchra in littoral mobile sand |
     Pontocrates arenarius in littoral mobile sand |
   Polychaete/amphipod-dominated fine sand shores |
    Polychaetes in littoral fine sand |
     Polychaetes, including Paraonis fulgens, in littoral fine sand |
     Polychaetes and Angulus tenuis in littoral fine sand |
     Nephtys cirrosa-dominated littoral fine sand |
   Polychaete/bivalve-dominated muddy sand shores |
    Macoma balthica and Arenicola marina in littoral muddy sand |
    Cerastoderma edule and polychaetes in littoral muddy sand |
    Hediste diversicolor, Macoma balthica and Eteone longa in littoral muddy sand |
    Bathyporeia pilosa and Corophium arenarium in littoral muddy sand |
    Lanice conchilega in littoral sand |
  Littoral mud |
   Polychaete/bivalve-dominated mid estuarine mud shores |
    Nephtys hombergii, Macoma balthica and Streblospio shrubsolii in littoral sandy mud |
    Hediste diversicolor and Macoma balthica in littoral sandy mud |
    Hediste diversicolor, Macoma balthica and Scrobicularia plana in littoral sandy mud |
   Polychaete/oligochaete-dominated upper estuarine mud shores |
    Nephtys hombergii and Streblospio shrubsolii in littoral mud |
    Hediste diversicolor in littoral mud |
     Hediste diversicolor and Streblospio shrubsolii in littoral sandy mud |
     Hediste diversicolor and Corophium volutator in littoral mud |
     Hediste diversicolor and oligochaetes in littoral mud |
    Tubificoides benedii and other oligochaetes in littoral mud |
  Littoral mixed sediment |
   Hediste-dominated gravelly sandy mud shores |
    Hediste diversicolor in littoral gravelly muddy sand and gravelly sandy mud |
     Hediste diversicolor and Macoma balthica in littoral gravelly mud |
     Hediste diversicolor and Scrobicularia plana in littoral gravelly mud |
     Hediste diversicolor and Streblospio shrubsolii in littoral gravelly sandy mud |
     Hediste diversicolor, cirratulids and Tubificoides spp. in littoral gravelly sandy mud |
     Hediste diversicolor and Corophium volutator in littoral gravelly sandy mud |
   Species-rich mixed sediment shores |
    Cirratulids and Cerastoderma edule in littoral mixed sediment |
  Littoral macrophyte-dominated sediment |
   Saltmarsh |
   Littoral seagrass beds |
    Zostera noltii beds in littoral muddy sand |
  Littoral biogenic reefs |
   Littoral Sabellaria honeycomb worm reefs |
    Sabellaria alveolata reefs on sand-abraded eulittoral rock |
   Littoral mussel beds on sediment |
   Mytilus edulis beds on littoral sediments |
   Mytilus edulis beds on littoral mixed substrata |
   Mytilus edulis beds on littoral sand |
   Mytilus edulis beds on littoral mud |
 Infralittoral rock (and other hard substrata) |
  High energy infralittoral rock |
   Kelp with cushion fauna and/or foliose red seaweeds |
    Alaria esculenta on exposed sublittoral fringe bedrock |
     Alaria esculenta, Mytilus edulis and coralline crusts on very exposed sublittoral fringe bedrock |
     Alaria esculenta and Laminaria digitata on exposed sublittoral fringe bedrock |
    Alaria esculenta forest with dense anemones and crustose sponges on extremely exposed infralittoral bedrock |
    Laminaria hyperborea forest with a faunal cushion (sponges and polyclinids) and foliose red seaweeds on very exposed upper infralittoral rock |
    Sparse Laminaria hyperborea and dense Paracentrotus lividus on exposed infralittoral limestone |
    Laminaria hyperborea with dense foliose red seaweeds on exposed infralittoral rock |
     Laminaria hyperborea forest with dense foliose red seaweeds on exposed upper infralittoral rock |
     Laminaria hyperborea park with dense foliose red seaweeds on exposed lower infralittoral rock |
     Mixed Laminaria hyperborea and Laminaria ochroleuca forest on exposed infralittoral rock |
    Foliose red seaweeds on exposed lower infralittoral rock |
     Foliose red seaweeds with dense Dictyota dichotoma and/or Dictyopteris membranacea on exposed lower infralittoral rock |
    Laminaria hyperborea and red seaweeds on exposed vertical rock |
   Sediment-affected or disturbed kelp and seaweed communities |
    Saccorhiza polyschides and other opportunistic kelps on disturbed sublittoral fringe rock |
    Laminaria saccharina and/or Saccorhiza polyschides on exposed infralittoral rock |
    Laminaria saccharina, Chorda filum and dense red seaweeds on shallow unstable infralittoral boulders or cobbles |
    Dense Desmarestia spp. with filamentous red seaweeds on exposed infralittoral cobbles, pebbles and bedrock |
    Mixed kelps with scour-tolerant and opportunistic foliose red seaweeds on scoured or sand-covered infralittoral rock |
    Halidrys siliquosa and mixed kelps on tide-swept infralittoral rock with coarse sediment |
    Polyides rotundus, Ahnfeltia plicata and Chondrus crispus on sand-covered infralittoral rock |
  Moderate energy infralittoral rock |
   Kelp and red seaweeds (moderate energy infralittoral rock) |
    Laminaria digitata on moderately exposed sublittoral fringe rock |
     Laminaria digitata on moderately exposed sublittoral fringe bedrock |
     Laminaria digitata and under-boulder fauna on sublittoral fringe boulders |
     Laminaria digitata and piddocks on sublittoral fringe soft rock |
    Laminaria hyperborea on tide-swept, infralittoral rock |
     Laminaria hyperborea forest, foliose red seaweeds and a diverse fauna on tide-swept upper infralittoral rock |
     Laminaria hyperborea park with hydroids, bryozoans and sponges on tide-swept lower infralittoral rock |
    Laminaria hyperborea on tide-swept infralittoral mixed substrata |
     Laminaria hyperborea forest and foliose red seaweeds on tide-swept upper infralittoral mixed substrata |
     Laminaria hyperborea park and foliose red seaweeds on tide-swept lower infralittoral mixed substrata |
    Laminaria hyperborea and foliose red seaweeds on moderately exposed infralittoral rock |
     Laminaria hyperborea forest and foliose red seaweeds on moderately exposed upper infralittoral rock |
     Laminaria hyperborea park and foliose red seaweeds on moderately exposed lower infralittoral rock |
     Grazed Laminaria hyperborea forest with coralline crusts on upper infralittoral rock |
     Grazed Laminaria hyperborea park with coralline crusts on lower infralittoral rock |
     Sabellaria spinulosa with kelp and red seaweeds on sand-influenced infralittoral rock |
    Dense foliose red seaweeds on silty moderately exposed infralittoral rock |
    Laminaria hyperborea on moderately exposed vertical rock. |
    Hiatella arctica and seaweeds on vertical limestone / chalk. |
   Kelp and seaweed communities in tide-swept sheltered conditions |
    Laminaria digitata, ascidians and bryozoans on tide-swept sublittoral fringe rock |
    Mixed kelp with foliose red seaweeds, sponges and ascidians on sheltered tide-swept infralittoral rock |
    Mixed kelp and red seaweeds on infralittoral boulders, cobbles and gravel in tidal rapids |
    Laminaria saccharina with foliose red seaweeds and ascidians on sheltered tide-swept infralittoral rock |
    Filamentous red seaweeds, sponges and Balanus crenatus on tide-swept variable-salinity infralittoral rock |
  Low energy infralittoral rock |
   Silted kelp communities (sheltered infralittoral rock) |
    Mixed Laminaria hyperborea and Laminaria ochroleuca forest on moderately exposed or sheltered infralittoral rock |
    Mixed Laminaria hyperborea and Laminaria saccharina on sheltered infralittoral rock |
     Mixed Laminaria hyperborea and Laminaria saccharina forest on sheltered upper infralittoral rock |
     Mixed Laminaria hyperborea and Laminaria saccharina park on sheltered lower infralittoral rock |
     Grazed, mixed Laminaria hyperborea and Laminaria saccharina on sheltered infralittoral rock |
    Laminaria saccharina on very sheltered infralittoral rock |
     Laminaria saccharina and Laminaria digitata on sheltered sublittoral fringe rock |
     Laminaria saccharina forest on very sheltered upper infralittoral rock |
     Laminaria saccharina park on very sheltered lower infralittoral rock |
     Grazed Laminaria saccharina with Echinus, brittlestars and coralline crusts on sheltered infralittoral rock |
    Silted cape-form Laminaria hyperborea on very sheltered infralittoral rock |
    Sargassum muticum on shallow slightly tide-swept infralittoral mixed substrata |
   Kelp in variable or reduced salinity |
    Codium spp. with red seaweeds and sparse Laminaria saccharina on shallow, heavily-silted, very sheltered infralittoral rock |
    Laminaria saccharina and Psammechinus miliaris on variable salinity grazed infralittoral rock |
    Laminaria saccharina with Phyllophora spp. and filamentous green seaweeds on variable or reduced salinity infralittoral rock |
   Faunal communities on variable or reduced salinity infralittoral rock |
    Mytilus edulis beds on reduced salinity infralittoral rock |
    Cordylophora caspia and Electra crustulenta on reduced salinity infralittoral rock |
    Hartlaubella gelatinosa and Conopeum reticulum on low salinity infralittoral mixed substrata |
   Submerged fucoids, green or red seaweeds (low salinity infralittoral rock) |
    Ascophyllum nodosum with epiphytic sponges and ascidians on variable salinity infralittoral rock |
    Mixed fucoids, Chorda filum and green seaweeds on reduced salinity infralittoral rock |
    Polyides rotundus and/or Furcellaria lumbricalis on reduced salinity infralittoral rock |
    Fucus ceranoides and Enteromorpha spp. on low salinity infralittoral rock |
  Features of infralittoral rock |
   Infralittoral surge gullies and caves |
    Foliose seaweeds and coralline crusts in surge gully entrances |
    Anemones, including Corynactis viridis, crustose sponges and colonial ascidians on very exposed or wave surged vertical infralittoral rock |
    Crustose sponges and colonial ascidians with Dendrodoa grossularia or barnacles on wave-surged infralittoral rock |
    Dendrodoa grossularia and Clathrina coriacea on wave-surged vertical infralittoral rock |
    Crustose sponges on extremely wave-surged infralittoral cave or gully walls |
    Coralline crusts in surge gullies and scoured infralittoral rock |
     Balanus crenatus and/or Pomatoceros triqueter with spirorbid worms and coralline crusts on severely-scoured vertical infralittoral rock |
     Coralline crusts and crustaceans on mobile boulders or cobbles in surge gullies |
   Infralittoral fouling seaweed communities |
 Circalittoral rock (and other hard substrata) |
  High energy circalittoral rock |
   Very tide-swept faunal communities |
    Balanus crenatus and Tubularia indivisa on extremely tide-swept circalittoral rock |
    Tubularia indivisa on tide-swept circalittoral rock |
     Tubularia indivisa and cushion sponges on tide-swept turbid circalittoral bedrock |
     Alcyonium digitatum with dense Tubularia indivisa and anemones on strongly tide-swept circalittoral rock |
   Deep sponge communities (circalittoral) |
    Phakellia ventilabrum and Axinellid sponges on deep, wave- exposed circalittoral rock |
   Mixed faunal turf communities |
    Bryozoan turf and erect sponges on tide-swept circalittoral rock |
     Eunicella verrucosa and Pentapora foliacea on wave-exposed circalittoral rock |
     Mixed turf of bryozoans and erect sponges with Dysidia fragilis and Actinothoe sphyrodeta on tide-swept wave-exposed circalittoral rock |
     Mixed turf of bryozoans and erect sponges with Sagartia elegans on tide-swept ciraclittoral rock |
    Corynactis viridis and a mixed turf of crisiids, Bugula, Scrupocellaria, and Cellaria on moderately tide-swept exposed circalittoral rock |
    Mixed turf of hydroids and large ascidians with Swiftia pallida and Caryophyllia smithii on weakly tide-swept circalittoral rock |
    Flustra foliacea and colonial ascidians on tide-swept moderately wave-exposed circalittoral rock |
     Polyclinum aurantium and Flustra foliacea on sand-scoured tide-swept moderately wave-exposed circalittoral rock |
     Flustra foliacea, small solitary and colonial ascidians on tide-swept circalittoral bedrock or boulders |
     Flustra foliacea and colonial ascidians on tide-swept exposed circalittoral mixed substrata |
    Sparse sponges, Nemertesia spp. and Alcyonidium diaphanum on circalittoral mixed substrata |
    Suberites spp. with a mixed turf of crisiids and Bugula spp. on heavily silted moderately wave-exposed shallow circalittoral rock |
    Flustra foliacea and Haliclona oculata with a rich faunal turf on tide-swept circalittoral mixed substrata |
    Molgula manhattensis with a hydroid and bryozoan turf on tide-swept moderately wave-exposed circalittoral rock |
    Sponges and anemones on vertical circalittoral bedrock |
  Moderate energy circalittoral rock |
   Echinoderms and crustose communities |
    Caryophyllia smithii and Swiftia pallida on circalittoral rock |
     Caryophyllia smithii, Swiftia pallida and Alcyonium glomeratum on wave-sheltered circalittoral rock |
     Caryophyllia smithii, Swiftia pallida and large solitary ascidians on exposed or moderately exposed circalittoral rock |
    Caryophyllia smithii, sponges and crustose communities on wave-exposed circalittoral rock |
     Brittlestars overlying coralline crusts, Parasmittina trispinosa and Caryophyllia smithii on wave-exposed circalittoral rock |
     Caryophyllia smithii and sponges with Pentapora foliacea, Porella compressa and crustose communities on wave-exposed circalittoral rock |
    Urticina felina and sand-tolerant fauna on sand-scoured or covered circalittoral rock |
    Faunal and algal crusts on exposed to moderately wave-exposed circalittoral rock |
     Flustra foliacea on slightly scoured silty circalittoral rock |
     Alcyonium digitatum, Pomatoceros triqueter, algal and bryozoan crusts on wave-exposed circalittoral rock |
     Alcyonium digitatum with Securiflustra securifrons on tide-swept moderately wave-exposed circalittoral rock |
     Brittlestars on faunal and algal encrusted exposed to moderately wave-exposed circalittoral rock |
     Faunal and algal crusts with Pomatoceros triqueter and sparse Alcyonium digitatum on exposed to moderately wave-exposed circalittoral rock |
     Caryophyllia smithii with faunal and algal crusts on moderately wave-exposed circalittoral rock |
    Alcyonium digitatum and faunal crust communities on vertical circalittoral bedrock |
   Circalittoral Sabellaria reefs (on rock) |
    Sabellaria spinulosa encrusted circalittoral rock |
     Sabellaria spinulosa with a bryozoan turf and barnacles on silty turbid circalittoral rock |
     Sabellaria spinulosa, didemnids and other small ascidians on tide-swept moderately wave-exposed circalittoral rock |
   Soft rock communities |
    Piddocks with a sparse associated fauna in sublittoral very soft chalk or clay |
    Polydora sp. tubes on moderately exposed sublittoral soft rock |
    Hiatella-bored vertical sublittoral limestone rock |
   Circalittoral mussel beds on rock |
    Mytilus edulis beds with hydroids and ascidians on tide-swept exposed to moderately wave-exposed circalittoral rock |
    Musculus discors beds on moderately exposed circalittoral rock |
   Circalittoral faunal communities in variable salinity |
    Cushion sponges and hydroids on turbid tide-swept sheltered circalittoral rock |
     Cushion sponges, hydroids and ascidians on turbid tide-swept sheltered circalittoral rock |
     Cushion sponges and hydroids on turbid tide-swept variable salinity sheltered circalittoral rock |
    Halichondria bowerbanki, Eudendrium arbusculum and Eucratea loricata on reduced salinity tide-swept circalittoral mixed substrata |
  Low energy circalittoral rock |
   Brachiopod and ascidian communities |
    Solitary ascidians, including Ascidia mentula and Ciona intestinalis, on wave-sheltered circalittoral rock |
     Solitary ascidians, including Ascidia mentula and Ciona intestinalis, with Antedon spp. on wave-sheltered circalittoral rock |
     Dense brittlestars with sparse Ascidia mentula and Ciona intestinalis on sheltered circalittoral mixed substrata |
    Large solitary ascidians and erect sponges on wave-sheltered circalittoral rock |
    Antedon spp., solitary ascidians and fine hydroids on sheltered circalittoral rock |
    Neocrania anomala and Protanthea simplex on sheltered circalittoral rock |
    Neocrania anomala and Protanthea simplex on very wave-sheltered circalittoral rock |
    Neocrania anomala, Dendrodoa grossularia and Sarcodictyon roseum on variable salinity circalittoral rock |
  Features of circalittoral rock |
   Circalittoral caves and overhangs |
    Sponges, cup corals and anthozoans on shaded or overhanging circalittoral rock |
   Circalittoral fouling faunal communities |
   Alcyonium digitatum and Metridium senile on moderately wave-exposed circalittoral steel wrecks |
   Ascidiella aspersa on circalittoral artificial substrata |
 Sublittoral sediment |
 Sublittoral coarse sediment (unstable cobbles and pebbles, gravels and coarse sands) |
  Sublittoral coarse sediment in variable salinity (estuaries) |
  Infralittoral coarse sediment |
   Sparse fauna on highly mobile sublittoral shingle (cobbles and pebbles) |
   Halcampa chrysanthellum and Edwardsia timida on sublittoral clean stone gravel |
   Moerella spp. with venerid bivalves in infralittoral gravelly sand |
   Hesionura elongata and Microphthalmus similis with other interstitial polychaetes in infralittoral mobile coarse sand |
   Glycera lapidum in impoverished infralittoral mobile gravel and sand |
   Cumaceans and Chaetozone setosa in infralittoral gravelly sand |
   Dense Lanice conchilega and other polychaetes in tide-swept infralittoral sand and mixed gravelly sand |
  Circalittoral coarse sediment |
   Pomatoceros triqueter with barnacles and bryozoan crusts on unstable circalittoral cobbles and pebbles |
   Mediomastus fragilis, Lumbrineris spp. and venerid bivalves in circalittoral coarse sand or gravel |
   Protodorvillea kefersteini and other polychaetes in impoverished circalittoral mixed gravelly sand |
   Neopentadactyla mixta in circalittoral shell gravel or coarse sand |
   Branchiostoma lanceolatum in circalittoral coarse sand with shell gravel |
  Offshore circalittoral coarse sediment |
   Glycera lapidum, Thyasira spp. and Amythasides macroglossus in offshore gravelly sand |
   Hesionura elongata and Protodorvillea kefersteini in offshore coarse sand |
 Sublittoral sands and muddy sands |
  Sublittoral sand in low or reduced salinity (lagoons) |
  Sublittoral sand in variable salinity (estuaries) |
   Infralittoral mobile sand in variable salinity (estuaries) |
   Nephtys cirrosa and Macoma balthica in variable salinity infralittoral mobile sand |
   Neomysis integer and Gammarus spp. in fluctuating low salinity infralittoral mobile sand |
  Infralittoral fine sand |
   Infralittoral mobile clean sand with sparse fauna |
   Sertularia cupressina and Hydrallmania falcata on tide-swept sublittoral sand with cobbles or pebbles. |
   Nephtys cirrosa and Bathyporeia spp. in infralittoral sand |
   Semi-permanent tube-building amphipods and polychaetes in sublittoral sand |
  Infralittoral muddy sand |
   Echinocardium cordatum and Ensis spp. in lower shore and shallow sublittoral slightly muddy fine sand |
   Fabulina fabula and Magelona mirabilis with venerid bivalves and amphipods in infralittoral compacted fine muddy sand |
   Arenicola marina in infralittoral fine sand or muddy sand |
   Spisula subtruncata and Nephtys hombergii in shallow muddy sand |
  Circalittoral fine sand |
   Echinocyamus pusillus, Ophelia borealis and Abra prismatica in circalittoral fine sand |
   Abra prismatica, Bathyporeia elegans and polychaetes in circalittoral fine sand |
  Circalittoral muddy sand |
   Abra alba and Nucula nitidosa in circalittoral muddy sand or slightly mixed sediment |
   Amphiura brachiata with Astropecten irregularis and other echinoderms in circalittoral muddy sand |
  Offshore circalittoral sand |
   Maldanid polychaetes and Eudorellopsis deformis in offshore circalittoral sand or muddy sand |
   Owenia fusiformis and Amphiura filiformis in offshore circalittoral sand or muddy sand |
 Sublittoral cohesive mud and sandy mud communities |
  Sublittoral mud in low or reduced salinity (lagoons) |
  Sublittoral mud in variable salinity (estuaries) |
   Polydora ciliata and Corophium volutator in variable salinity infralittoral firm mud or clay |
   Aphelochaeta marioni and Tubificoides spp. in variable salinity infralittoral mud |
   Nephtys hombergii and Tubificoides spp. in variable salinity infralittoral soft mud |
   Infralittoral fluid mobile mud |
   Capitella capitata and Tubificoides spp. in reduced salinity infralittoral muddy sediment |
   Oligochaetes in variable or reduced salinity infralittoral muddy sediment |
   Limnodrilus hoffmeisteri, Tubifex tubifex and Gammarus spp. in low salinity infralittoral muddy sediment |
  Infralittoral sandy mud |
   Nephtys hombergii and Macoma balthica in infralittoral sandy mud |
   Sagartiogeton undatus and Ascidiella aspersa on infralittoral sandy mud |
   Mysella bidentata and Abra spp. in infralittoral sandy mud |
   Melinna palmata with Magelona spp. and Thyasira spp. in infralittoral sandy mud |
   Ampelisca spp., Photis longicaudata and other tube-building amphipods and polychaetes in infralittoral sandy mud |
   Capitella capitata in enriched sublittoral muddy sediments |
  Infralittoral fine mud |
   Cerastoderma edule with Abra nitida in infralittoral mud |
   Arenicola marina in infralittoral mud |
   Philine aperta and Virgularia mirabilis in soft stable infralittoral mud |
   Ocnus planci aggregations on sheltered sublittoral muddy sediment |
   Beggiatoa spp. on anoxic sublittoral mud |
  Circalittoral sandy mud |
   Amphiura filiformis, Mysella bidentata and Abra nitida in circalittoral sandy mud |
   Thyasira spp. and Nuculoma tenuis in circalittoral sandy mud |
   Amphiura filiformis and Nuculoma tenuis in circalittoral and offshore sandy mud |
   Virgularia mirabilis and Ophiura spp. with Pecten maximus on circalittoral sandy or shelly mud |
    Virgularia mirabilis and Ophiura spp. with Pecten maximus, hydroids and ascidians on circalittoral sandy or shelly mud with stones |
   Lagis koreni and Phaxas pellucidus in circalittoral sandy mud |
  Circalittoral fine mud |
   Seapens and burrowing megafauna in circalittoral fine mud |
    Seapens, including Funiculina quadrangularis, and burrowing megafauna in undisturbed circalittoral fine mud |
   Burrowing megafauna and Maxmuelleria lankesteri in circalittoral mud |
   Brissopsis lyrifera and Amphiura chiajei in circalittoral mud |
  Offshore circalittoral mud |
   Ampharete falcata turf with Parvicardium ovale on cohesive muddy sediment near margins of deep stratified seas |
   Foraminiferans and Thyasira sp. in deep circalittoral fine mud |
   Styela gelatinosa, Pseudamussium septemradiatum and solitary ascidians on sheltered deep circalittoral muddy sediment |
   Capitella capitata and Thyasira spp. in organically-enriched offshore circalittoral mud and sandy mud |
    Capitella capitata, Thyasira spp. and Ophryotrocha dubia in organically-enriched offshore circalittoral sandy mud |
   Levinsenia gracilis and Heteromastus filifirmis in offshore circalittoral mud and sandy mud |
   Paramphinome jeffreysii, Thyasira spp. and Amphiura filiformis in offshore circalittoral sandy mud |
   Myrtea spinifera and polychaetes in offshore circalittoral sandy mud |
 Sublittoral mixed sediment |
  Sublittoral mixed sediment in low or reduced salinity (lagoons) |
  Sublittoral mixed sediment in variable salinity (estuaries) |
   Aphelochaeta spp. and Polydora spp. in variable salinity infralittoral mixed sediment |
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