Layout and components of biotope
descriptions
Descriptions for each unit in the classification, from broad
habitats to sub-biotopes, are laid out as follows:
Code
A unique letter code, reflecting the level of the described
type within the classification hierarchy. A "breadcrumb trail" of
codes is shown in the standard classificaiton colour scheme, which
indicates what level of the biotope classification is being
currently looked at, and allows a direct link to the "parents" of
the type currently being looked at.
Title
The title gives the key biological and physical features of
the type, with emphasis on the features which help to distinguish
it from closely related types of the same level in the hierarchy.
The habitat part of the title usually includes the zone, substratum
and another key habitat factor. To avoid becoming overly clumsy the
titles do not cover all habitat characteristics or
characterising species, and common names are not given (although
they are given in the text description).
NOTE: It is very important to refer to the full
description and to the habitat matrices to determine the full
nature of the type and not to rely on the title
alone.
Habitat
characteristics
The typical habitat characteristics of the type for salinity,
wave exposure, tidal currents, substratum, zone, height or depth
band and, where appropriate, other factors critical to that
particular type. The range given for each factor tends to be
broader for higher types and more tightly defined for lower types.
When assigning samples to types, it should be noted that in some
cases the type may occur outside the range given (see profiles
given in the
comparative tables which show that a small
proportion of records may occur outside the typical range for the
type), though care should be taken to ensure that another type has
not been described to cover the example being considered. All
heights and depths are corrected to chart datum.
Previous
code
Codes used in versions 6.95, 96.7, and 97.06 (Connor
et
al. 1995 a, 1996, 1997 a, b) are given where different to the
current code. Where communities from previous versions have been
combined or split, previous codes are shown as far as possible.
Some communities in the revised classification are newly defined
and may not relate directly to types in the previous
classification. Lookup tables which help to translate 2004 to 1997
codes (and vice versa) are available to download in the form of an
Excel™ spreadsheet from the
main download page.
A distribution map is included, showing the location of field
survey records assigned to the biotope on the JNCC marine database.
The red dots show the location of core biotope records, i.e. those
records on which the biotope description is based (see
data analysis for
details). Blue dots show other certain records of the biotope,
whilst black dots denote uncertain records of that biotope (i.e.
field records that have been tentatively assigned to the biotope -
they may not match the description fully, or they may be
incomplete records). Note that the distribution
maps only show those records that are held on
the JNCC marine database, and therefore do not reflect the
full extent of the biotope.
Description
An account of the general nature of the habitat and community
characteristics, and its micro-habitat features (e.g. crevices,
under-boulders, kelp stipes) if present.
Situation
Describes the general situation on the shore or in the
sublittoral, in relation to other types (i.e. along gradients of
substratum, zonation, wave exposure, tidal currents, salinity
etc.).
Temporal
variation
This section outlines the known natural temporal dynamics of
the type described, such as seasonal changes in community structure
or physical environment. In general, much more information is
needed for this section. In some cases separate types may have been
defined because there is a lack of knowledge that the communities
are temporal variations within a single habitat type.
Similar
types
Attention is drawn to similar types which should be considered
before assigning a field record to a particular biotope. The main
similarities and principal distinguishing features are described
for each similar biotope, and the codes are hyperlinked directly to
the relevant descriptions.
Characterising
species
A list of those species which contribute most to the overall
similarity between core records assigned to the type, i.e.
characterise the type, with associated information on their
frequency of occurrence, their individual contribution to the
similarity within the core data set of records, and the typical
abundance at which they occur.
For each type, characterising species have been determined
using the SIMPER routine in PRIMER (Clarke & Warwick, 2001).
For a given set of records (in this case, core records of each
type), SIMPER indicates and ranks the individual contribution of
each species to the overall similarity within the data set. Both
the frequency of occurrence of each species within the dataset and
their abundance (using the
SACFOR abundance scale) for epifaunal data and
numeric counts for infaunal data) are taken into account during
this process. Species that contribute more than 1% to the
overall similarity of the records within the data set are defined
as 'characterising species', and listed in a characterising species
table. Those that contribute less that 1% are not listed. Species
which qualify according to the SIMPER routine, but are Present or
Rare on the MNCR SACFOR scale and present in fewer than 20% of the
records, are occasionally excluded from the characterising species
table.
Care has been taken to mention each of the characterising
species in the descriptions for each type. Sometimes additional
species are mentioned that are particularly indicative (faithful)
of that type or characteristic of a biogeographic region, but which
have not qualified as 'characterising species' according to the
SIMPER routine.
Some of the biotope descriptions, especially in the
sublittoral sediment section, have been based on a mixture of
epifaunal (semi-quantitative) and infaunal (quantitative) sample
data. In these cases, separate SIMPER analyses were carried out for
the two types of data, and the outcome was combined into a single
characterising species table. Where there is overlap between
species recorded in the epifaunal and infaunal data, there are
duplicate entries for species: the entries relating to the infaunal
datasets have figures for "numbers per metres squared", whereas
entries relating to epifaunal datasets only have SACFOR
entries.
The % contribution to similarity column
of the table shows the contribution of each characterising species
to the similarity within the type, i.e. the higher the
contribution, the higher the importance of the species. The number
of species in the table reflects the species diversity within each
type. In types with a high species richness, a large number of
species each contribute with a relatively low amount to the
similarity within the group. If a type has low diversity, then a
small number of species contribute with relatively large amounts to
the overall similarity and hence fewer species are listed in the
table. In a few cases, a long species list indicates low overall
similarity of records within the type.
The % frequency of occurrence column of
the table shows the occurrence of a species within a certain
biotope. The symbols represent percentage occurrence in the samples
as follows:
Occurs in 81-100% of the records for the type
Occurs in 61-80% of the records for the type
Occurs
in 41-60% of the records for the type
Occurs
in 21-40% of the records for the type
The typical abundance column of the table
shows the mean SACFOR abundance for each characterising species
within the samples where it is present. Quantitative infaunal
counts have been converted to the SACFOR scale for compatibility of
data presentation. For types where the core records are exclusively
quantitative infaunal records (e.g. most of the littoral sediment
types), an additional column is included in the characterising
species table, showing mean counts per m2 for each
species within the core data set.
Example photographs
Where they are available, up to three photographs are shown to
illustrate the appearance of the biotope in the field.