These habitats occur above the high water mark, but often in
areas which are influenced by wave splash and sea-spray.
Salt-tolerant species are the characteristic colonisers of this
habitat and the biotopes present are strongly influenced by
sediment size as well as the degree of wave exposure of the
shore.
Strandline communities are often present on moderately exposed
sandy shores, particularly on flat, slightly mobile beaches with
little or no human disturbance. Under these conditions annual
vegetation can develop on the accumulations of drift material rich
in nitrogenous organic matter at or near the high water mark.
Characteristic vascular plants include sea sandwort
Honckenya peploides, saltwort Salsola kali, and
sea beet Beta maritima.
On the upper margins of the shore, three major supralittoral
sediment habitat types occur: coastal vegetated shingle, sand
dunes, and machair.
Shingle beaches are widely distributed around the UK coastline
and tend to form in high-energy environments where the sea can pile
up pebbles above the tide line. There are five recognised
types distributed around the UK: fringing beaches, spits, barriers,
cuspate forelands and barrier islands. Herb-rich open pioneer
stages colonise the seaward edge with species such as sea-kale
Crambe maritima, sea pea Lathyrus japonicus,
thrift Armeria maritima, yellow horned-poppy Glaucium
flavum and sea holly Eryngium maritimum.
Grassland, heath, scrub, and moss and lichen-dominated vegetation
of old, stable, shingle occur further inland.
Distinct features within sand dune systems include fore dunes,
yellow dunes, dune grassland, dune slacks, dune heath and dune
scrub. Factors such as stability and moisture retention in
these different systems determine what species are present.
In mobile fore dunes, for example, typical species are
marram Ammophila arenaria and, in northern areas of
Britain, lyme-grass Leymus arenarius. Dune slacks,
which are areas of wetland within the dune system, may have scarce
plants such as fen orchid Liparis loeselii
and petalwort Petalophyllum ralfsii.
Machair is a distinctive sand dune formation that is only
found on the North and West coast of Scotland and in western
Ireland. The soils are made up of wind deposited shell-sand
blown inland from coastal beaches and mobile dunes, which lie over
impermeable rock. The main habitats of machair are dry
grassland, damp grassland, marsh and standing water and the
vegetation is broadly described as a herb-rich sward. Machair
also supports a rich invertebrate fauna and large wader
populations, for example on the Uists, Tiree and Coll. These
populations of waders are considered the most important in the
north-west Palaearctic.